INDIANAPOLIS — The Indianapolis Colts’ pass rush was shut out Sunday – no sacks and no official quarterback hits.

And the Jacksonville Jaguars used a very familiar tactic to help keep quarterback Trevor Lawrence clean – the quick passing game.

In seven career games against the Colts, Lawrence has completed 72.6% of his passes and has a quarterback rating of 104.4. He was 28-of-34 (82.4%) in a 37-34 Jaguars victory Sunday with a career-high 371 yards.

“We never got them really off schedule,” Indianapolis defensive coordinator Gus Bradley said Tuesday. “There were times where those tackling (issues) take place if you look. I mean, we had a chance to get them second-and-9. They fell forward. They got 5 extra yards. So, we never really got them into those situations where it was a drop-back pass.”

Missed tackles have been a recurring problem this season.

According to Pro Football Focus, the Colts are tied for third in the NFL with 60 missed tackles through the first five games. Only the Las Vegas Raiders (66) and Atlanta Falcons (62) have more.

That’s been a major contributing factor for a defense that ranks last in the 32-team league in total yards allowed (2,096), 31st in first downs surrendered (31) and 30th in average yards per pass attempt (7.4).

“I think a lot of times it’s like we make contact, but we’re not finishing,” Bradley said. “We always say you’ve got to come up with a body part, come up with something – wrap up and drive for five. I just don’t think we’re finishing like we need to, especially in space. Space tackling is where you’re seeing some of the issues occur.”

As Indianapolis (2-3) prepares for another matchup with an AFC South rival this week on the road against the Tennessee Titans (1-3), Bradley is under heavy fire.

The defense ranks 25th with an average of 24.4 points allowed per game and surrendered 17 points in the fourth quarter alone against Jacksonville last week.

Head coach Shane Steichen has repeatedly expressed his faith and confidence in Bradley, but the veteran coordinator admits he hears the outside criticism.

“It is difficult,” Bradley said. “I mean, I think you look more at it as for the organization, for the players, for the team, for the fans – that’s the part that is difficult. You want to play better. Our whole emphasis is on getting better each week. There was a period of time last game where we weren’t getting better. We didn’t get better at tackling. We didn’t get better at some of our fundamental things, and it showed.

“That’s the disappointment is that we had an opportunity to win a game, and we didn’t pull our end of the bargain – through explosive plays, not making as many plays as we could have — and that’s extremely disappointing. But, yeah, it’s tough. It’s always humbling when you go through a game like that. You feel like the defense had a big part of it, and you didn’t show up the way you should have.”

ROSTER MOVES

The Colts signed cornerback David Long Jr. to the 53-man roster from the practice squad Tuesday and re-signed running back Trent Pennix to the practice squad.

A third-round pick out of Michigan in 2019, Long has appeared in 68 games with 12 starts for the Los Angeles Rams, Las Vegas Raiders, Carolina Panthers, Green Bay Packers and Indianapolis.

Long appeared in two games last season with the Colts, playing 12 snaps on special teams. He has 96 career tackles, eight pass breakups and one interception.

Indianapolis still has one open spot on the 53-man roster.